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List of works by Johan Billen

A Combination of Fertility Signals and Aggression Regulates Reproduction in the Ant Gnamptogenys striatula

article by Els Lommelen et al published 10 April 2010 in Journal of Insect Behavior

A new type of exocrine gland and its function in mass recruitment in the ant Cylindromyrmex whymperi (Formicidae, Cerapachyinae) ⬇️

scientific article published on September 1, 2001

A novel exocrine structure of the bicellular unit type in the thorax of termites

scientific article published in 2010

A novel mechanism for jumping in the indian antHarpegnathos saltator (Jerdon) (Formicidae, Ponerinae)

Aberrant Collagen Composition of the Trabecular Meshwork Results in Reduced Aqueous Humor Drainage and Elevated IOP in MMP-9 Null Mice

scientific article

Age-dependent morphology and ultrastructure of the cornua glands in drones of<i>Apis mellifera</i>

scientific article published in 2009

Amfor expression in the honeybee brain: a trigger mechanism for nurse-forager transition

scientific article published on 3 August 2008

An overlooked mandibular-rubbing behavior used during recruitment by the African weaver ant, Oecophylla longinoda.

scientific article

Behavioral mechanisms and morphological symptoms of zombie ants dying from fungal infection

scientific article

Caste fate conflict in swarm-founding social hymenoptera: an inclusive fitness analysis.

scientific article published in July 2003

Chemistry of metapleural gland secretions of three attine ants,Atta sexdens rubropilosa, Atta cephalotes, andAcromyrmex octospinosus (Hymenoptera: Formicidae).

scientific article

Cloning and expression of PKG, a candidate foraging regulating gene in Vespula vulgaris

article

Cloning and sequencing of wsp encoding gene fragments reveals a diversity of co-infecting Wolbachia strains in Acromyrmex leafcutter ants

scientific article

Co-occurrence of three types of egg policing in the Norwegian wasp Dolichovespula norwegica

article by Wim Bonckaert et al published 24 September 2010 in Behavioral Ecology and Sociobiology

Coevolved crypts and exocrine glands support mutualistic bacteria in fungus-growing ants

scientific article

Colony composition and behavioral characteristics of Myrmoteras iriodum and M. jaitrongi in Ulu Gombak, Peninsular Malaysia (Hymenoptera: Formicidae) ⬇️

scholarly article published in 2017

Colony stage and not facultative policing explains pattern of worker reproduction in the Saxon wasp

scientific article published on 18 July 2011

Comparative analysis of worker head anatomy of Formica and Brachyponera (Hymenoptera: Formicidae) ⬇️

scientific article published in 2020

Comparative morphology of cephalic exocrine glands among castes of the black ant Lasius niger

scientific article published on 30 January 2007

Comparative survey of abdominal gland secretions of the ant subfamily Ponerinae

scientific article published in January 2003

Comparison of spermatheca morphology between reproductive and non-reproductive females in social wasps

scientific article published on 19 November 2007

Conserved class of queen pheromones stops social insect workers from reproducing

scientific article

Contents of dufour glands of workers of three species of Tetramorium (Hymenoptera: Formicidae)

scientific article published on 01 March 1986

Cuticular hydrocarbons provide reliable cues of fertility in the ant Gnamptogenys striatula.

scientific article published on 11 August 2006

Defensive function of petiole spines in queens and workers of the formicine ant Polyrhachis lamellidens (Hymenoptera: Formicidae) against an ant predator, the Japanese tree frog Hyla japonica ⬇️

scholarly article published in 2016

Degeneration of sperm reservoir and the loss of mating ability in worker ants

scientific article published on 13 August 2008

Degeneration patterns of the worker spermatheca during morphogenesis in ants (Hymenoptera: Formicidae).

scientific article

Deleterious Wolbachia in the ant Formica truncorum.

scientific article published in March 2002

Development of the labial gland of the ponerine ant Pachycondyla obscuricornis (Hymenoptera, Formicidae) during the pupal stage

scientific article published on 01 October 2003

Differences in Climbing Ability of Cimex lectularius and Cimex hemipterus (Hemiptera: Cimicidae).

scientific article

Distinctive fungal communities in an obligate African ant-plant mutualism.

scientific article published in March 2017

Distribution and comparative morphology of the cloacal gland in ants (Hymenoptera : Formicidae)

scholarly article by Tom Wenseleers et al published April 1998 in Arthropod Structure and Development

Diversity and morphology of abdominal glands in workers of the ant genus Myopias (Formicidae, Ponerinae).

scientific article published on 18 January 2013

Dufour gland contents of three species ofMyrmecia (Hymenoptera: Formicidae), primitive ants of australia

scientific article published in August 1989

Effects of sublethal doses of crop protection agents on honey bee (Apis mellifera) global colony vitality and its potential link with aberrant foraging activity.

scientific article published in January 2009

Effects of the Argentine ant venom on terrestrial amphibians

scientific article published on 18 August 2020

Egg marking in the facultatively queenless ant Gnamptogenys striatula: the source and mechanism

scientific article published on 15 February 2008

Evolution of a soldier caste specialized to lay unfertilized eggs in the ant genus Crematogaster (subgenus Orthocrema) ⬇️

scientific article published on February 28, 2013

Evolution of cheaper workers in ants: a comparative study of exoskeleton thickness

scientific article published in 2017

Evolution of specialized spermatheca morphology in ant queens: insight from comparative developmental biology between ants and polistine wasps

scientific article published on 15 September 2009

Exocrine Glands and Their Key Function in the Communication System of Social Insects ⬇️

scholarly article in Formosan Entomologist, vol. 31 no. 2, 2011

Exocrine chemistry of the myrmicine ant Zacryptocerus pusillus (Hymenoptera: Formicidae).

scientific article published in September 2006

Exocrine glands in the legs of the social wasp Vespula vulgaris

scientific article published on 8 September 2015

Exocrine glands in the legs of the stingless bee Frieseomelitta varia (Lepeletier) (Apidae: Meliponini)

scientific article published on 21 January 2015

Extracellular vesicles spread the RNA interference signal of Tribolium castaneum TcA cells

scientific article published on 14 April 2020

Flight muscle histolysis in Lasius niger queens ⬇️

scholarly article published in 2021

Functional analysis of a pancreatic secretory trypsin inhibitor-like protein in insects: silencing effects resemble the human pancreatic autodigestion phenotype

scientific article published on 8 May 2011

Functional morphology of the maxillary and propharyngeal glands of Monomorium pharaonis (L.).

scientific article

Glandular innovations for a tunnelling life: Silk and associated leg glands in Melissotarsus and Rhopalomastix queen and worker ants

scientific article published on 17 August 2020

Histological study of the spermatheca in three thelytokous parthenogenetic ant species, Pristomyrmex punctatus, Pyramica membranifera and Monomorium triviale (Hymenoptera: Formicidae)

scholarly article by Ayako Gotoh et al published 18 February 2011 in Acta Zoologica

Histology of structures used in territorial combat by Borneo’s ‘exploding ants’

scientific article published in 2011

Hydrocarbon signatures of egg maternity, caste membership and reproductive status in the common wasp.

scientific article published on 11 January 2012

Insect exocrine glands

scientific article published on 02 September 2015

Intraspecific queen parasitism in a highly eusocial bee.

scientific article

Large interclone differences in melezitose secretion in the facultatively ant-tended black bean aphid Aphis fabae ⬇️

scientific article published on August 26, 2011

Levels of clonal mixing in the black bean aphid Aphis fabae, a facultative ant mutualist.

scientific article published on 21 July 2011

Linking the Morphology of Sternal Glands to Rubbing Behavior by <i>Vespa soror</i> (Hymenoptera: Vespidae) Workers During Recruitment for Group Predation

scientific article published in 2021

Metformin promotes lifespan through mitohormesis via the peroxiredoxin PRDX-2

scientific article

Morpho-functional comparison of the Dufour gland in the female castes of the Amazon ant Polyergus rufescens (Hymenoptera, Formicidae)

scholarly article by Donato A. Grasso et al published 28 July 2005 in Zoomorphology

Morphological and ultrastructural changes in the mucus glands of <i>Apis mellifera</i> drones during pupal development and sexual maturation

scientific article published in 2005

Morphology and ultrastructure of Dufour's and venom glands in the ant Camponotus japonicus Mayr (Hymenoptera: Formicidae).

scientific article published on 31 October 2017

Morphology and ultrastructure of Pavan’s gland of Aneuretus simoni (Formicidae, Aneuretinae) ⬇️

scholarly article published in 2016

Morphology and ultrastructure of a specialized bacterial pouch in the digestive tract of Tetraponera ants (Formicidae, Pseudomyrmecinae)

scientific article published on 01 July 2000

Morphology and ultrastructure of the Dufour gland in workers of social wasps (Hymenoptera, Vespidae).

scientific article

Morphology and ultrastructure of the Dufour gland of Myzinum sp. (Tiphiidae)

scientific article published on 28 April 2017

Morphology and ultrastructure of the infrabuccal pocket and its lining epithelium in workers of Ectomomyrmex javanus (Hymenoptera: Formicidae)

scientific article published on 05 September 2018

Morphology and ultrastructure of the mandibular gland in the ant Brachyponera sennaarensis (Hymenoptera, Formicidae).

scientific article

Morphology and ultrastructure of the venom gland in the ant Brachyponera sennaarensis ⬇️

scholarly article published in 2018

Morphology of a novel glandular epithelium lining the infrabuccal cavity in the ant Monomorium pharaonis (Hymenoptera, Formicidae)

scientific article published on 01 October 2004

Morphology of the Dufour gland within the honey bee sting gland complex

scientific article published in 2005

Morphology of the cervical lobe of the endophallus in<b><i>Apis mellifera</i></b>drones

scientific article published in 2007

NOTE: FINE STRUCTURE OF THE SECRETORY TUBULES OF THE VENOM GLAND IN THE EUMENID WASP RHYNCHIUM CYANOPTERUM ⬇️

scientific article

Novel exocrine glands in the foreleg coxae of Discothyrea ants

scientific article published on 17 August 2020

Novel thoracic glands in the ant Myopias hollandi

scientific article published on 23 April 2018

Occurrence and structural organization of the exocrine glands in the legs of ants.

scientific article published on October 2008

Ontogeny and morphology of the bulbus, part of the male reproductive organ in Apis mellifera carnica (Hymenoptera, Apidae)

scientific article published in 2011

Pleiotropic and novel phenotypes in the Drosophila gut caused by mutation of drop-dead.

scientific article published on 22 January 2018

Policing behaviour towards virgin egg layers in a polygynous ponerine ant.

scientific article

Pretarsus structure in relation to climbing ability in the ants Brachyponera sennaarensis and Daceton armigerum

scientific article

Queen-worker differences in spermatheca reservoir of phylogenetically basal ants

scientific article

Skeletomuscular adaptations of head and legs of ants for tunnelling through living wood

scientific article published on 14 August 2018

Sneaky queens in Melipona bees selectively detect and infiltrate queenless colonies

Social insect histology from the nineteenth century: the magnificent pioneer sections of Charles Janet

scientific article published on 15 July 2007

Spectacular Batesian mimicry in ants

scientific article published in October 2004

Structural adaptations and mechanism of reflex bleeding in the larvae of the myrmecophilous ladybird Diomus thoracicus

scientific article

Structural and physical evidence for an endocuticular gold reflector in the tortoise beetle, Charidotella ambita

scientific article

Structure and chemistry of the Dufour gland inPristomyrmexants (Hymenoptera, Formicidae)

scholarly article by Johan Billen et al published April 2000 in Acta Zoologica

Sublethal effects of crop protection on honey bee pollination: foraging behaviour and flower visits.

scientific article published in January 2008

Successful maintenance of a stingless bee population despite a severe genetic bottleneck

article published in 2010

Survey of the exocrine system in Protanilla wallacei (Hymenoptera, Formicidae).

scientific article published on 16 January 2013

Task-related variation of postpharyngeal and cuticular hydrocarbon compositions in the ant Myrmicaria eumenoides

scientific article published on 01 October 2000

Tetraponera ants have gut symbionts related to nitrogen-fixing root-nodule bacteria.

scientific article

The basitarsal sulcus gland, a novel exocrine structure in ants

The diversity of microorganisms associated with Acromyrmex leafcutter ants

scientific article

The dufour gland and the secretion placed on eggs of two species of social wasps,Liostenogaster flavolineata andParischnogaster jacobsoni (vespidae: Stenogastrinae)

scientific article published on 01 February 1993

The effects of four crop protection products on the morphology and ultrastructure of the hypopharyngeal gland of the European honeybee, Apis mellifera

scientific article published in 2011

The evolution of worker–queen polymorphism in Cataglyphis ants: interplay between individual- and colony-level selections

article by Fernando Amor et al published 4 March 2011 in Behavioral Ecology and Sociobiology

The exocrine system of Aneuretus simoni (Formicidae, Aneuretinae) ⬇️

scholarly article published in 2017

The importance of the spermathecal duct in bumblebees

scientific article published in September 2000

The intramandibular gland of Aneuretus simoni (Formicidae, Aneuretinae) ⬇️

scholarly article published in 2016

The life of a dead ant: the expression of an adaptive extended phenotype

scientific article

The metapleural gland of Aneuretus simoni (Formicidae, Aneuretinae) ⬇️

scholarly article published in 2017

The queen is dead--long live the workers: intraspecific parasitism by workers in the stingless bee Melipona scutellaris

scientific article published on 10 September 2009

The sting bulb gland in myrmecia and Nothomyrmecia (Hymenoptera : Formicidae): A new exocrine gland in ants

scientific article (publication date: 1990)

Thelytokous parthenogenesis by queens in the dacetine ant Pyramica membranifera (Hymenoptera: Formicidae).

scientific article

Ultrastructure of the mandibular gland of the ant Myrmoteras iriodum

scientific article

Uncovering head gland diversity in neotropical Polistinae wasps (Hymenoptera, Vespidae): comparative analysis and description of new glands

scientific article published on 23 June 2015

Variation with caste of the mandibular gland secretion in the leaf-cutting antAtta sexdens rubropilosa

scientific article published on 01 May 1993

Vestigial spermatheca morphology in honeybee workers, Apis cerana and Apis mellifera, from Japan

scientific article published in 2012

Vibrational communication between a myrmecophilous butterfly Spindasis lohita (Lepidoptera: Lycaenidae) and its host ant Crematogaster rogenhoferi (Hymenoptera: Formicidae)

scientific article published on 06 December 2019

Volatile glandular secretions of three species of new world army ants,Eciton burchelli, Labidus coecus, andLabidus praedator

scientific article published on 01 November 1993

Volatile secretion of dufour gland of workers of an army ant,Dorylus (Anomma) molestus.

scientific article published in August 1991

Volatile secretions of old world army antAenictus rotundatus and chemotaxonomic implications of army ant dufour gland chemistry

scientific article

Wax gland size according to worker age in Friesella schrottkyi

scientific article published in 2018

Wax glands of the horned gall aphid, Schlechtendalia chinensis, at different stages

scientific article published on 10 August 2020

Widespread occurrence of the micro-organism Wolbachia in ants

scientific article published in August 1998

Worker reproduction of the invasive yellow crazy ant Anoplolepis gracilipes ⬇️

scientific article (publication date: 2017)

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