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Authors whose works are in public domain in at least one jurisdiction

List of works by Alan Grafen

A first formal link between the price equation and an optimization program.

scientific article published in July 2002

A new model for discrete character evolution.

scientific article published on January 1997

Altruism via kin-selection strategies that rely on arbitrary tags with which they coevolve

scientific article published in August 2004

An inclusive fitness analysis of altruism on a cyclical network

scientific article published on 01 November 2007

Biological fitness and the Price Equation in class-structured populations

scientific article published on 19 February 2015

Biological fitness and the fundamental theorem of natural selection

scientific article published on 15 May 2015

Biological signals as handicaps

scientific article

Changes in parasite aggregation with age: a discrete infection model

Colony-level sex ratio selection in the eusocial Hymenoptera

scholarly article by J. J. Boomsma & A. Grafen published May 1991 in Journal of Evolutionary Biology

Defining fitness in an uncertain world

scientific article

Detecting kin selection at work using inclusive fitness

scientific article published in March 2007

Does the negative binomial distribution add up?

scientific article published on 01 December 1993

Error-prone signalling

scientific article published on 01 June 1992

Evolutionarily stable nesting strategy in a digger wasp

scientific article published on April 21, 1979

Fertility and labour supply in Femina economica

scientific article published on 01 October 1998

Fisher the evolutionary biologist

Formal Darwinism, the individual-as-maximizing-agent analogy and bet-hedging

scientific article published in 1999

Formalizing Darwinism and inclusive fitness theory.

scientific article published on November 2009

Foundations of a mathematical theory of darwinism.

scientific article published on 9 July 2013

Genetic scrambling as a defence against meiotic drive.

scientific article published on December 1991

Green beard as death warrant

scientific article published in Nature

How not to measure inclusive fitness.

scientific article published in July 1982

INTRASPECIFIC VARIATION IN ANT SEX RATIOS AND THE TRIVERS-HARE HYPOTHESIS.

scientific article published on July 1990

Inclusive fitness theory and eusociality

scientific article published on March 2011

Intraspecific Variation in Ant sex Ratios and the Trivers-Hare Hypothesis

Life history correlations and demography

scientific article published in Nature

Mate conflict and male behaviour in a solitary wasp, Trypoxylon (Trypargilum) politum (Hymenoptera: Sphecidae)

scholarly article by H.Jane Brockmann & Alan Grafen published February 1989 in Animal Behaviour

Natural selection of altruism in inelastic viscous homogeneous populations

scientific article published on 31 January 2008

Of mice and the MHC

scientific article published in Nature

Optimization of inclusive fitness.

scientific article published on 25 July 2005

Pheromones, social behaviour and the functions of secondary metabolism in bacteria

scientific article published on 01 March 1995

Relatedness with different interaction configurations

scientific article published on 13 October 2009

Richard Dawkins: How a Scientist Changed the Way We Think

biography and festschrift for biologist Richard Dawkins

Sex and flagellation.

scientific article

Sexual selection unhandicapped by the Fisher process.

scientific article published in June 1990

The Phylogenetic Regression

scientific article (publication date: 21 December 1989)

The Price equation and reproductive value

scientific article published on 09 March 2020

The continuous Sir Philip Sidney game: A simple model of biological signalling

scientific article published on May 21, 1992

The evolution of index signals to avoid the cost of dishonesty

scientific article published in September 2014

The formal Darwinism project: a mid-term report.

scientific article published on July 2007

The illusion of invariant quantities in life histories

scientific article

The left hand side of the Fundamental Theorem of Natural Selection

scientific article published on 20 July 2018

The simplest formal argument for fitness optimization.

scientific article published on December 2008

Total reproductive values for females and for males in sexual diploids are not equal

scientific article published on 21 May 2014

Unusual mutational mechanisms and evolution

scientific article

Various remarks on Lehmann and Keller's article

scientific article published on 01 September 2006

Why we need ESS signalling theory

scientific article published on 01 May 1993