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List of works by Keith Richard Willmott

A Comprehensive and Dated Phylogenomic Analysis of Butterflies

scientific article

A New Species Of Actinote Hübner from the Eastern Andes of Ecuador (Lepidoptera: Nymphalidae: Heliconiinae)

scholarly article by Keith Richard Willmott et al published January 2009 in Proceedings of the Entomological Society of Washington

A Review of Colobura (Lepidoptera: Nymphalidae) with Comments on Larval and Adult Ecology and Description of a Sibling Species

scientific article published in March 2001

A global phylogeny of butterflies reveals their evolutionary history, ancestral hosts and biogeographic origins

scientific article published in 2023

A new euptychiine butterfly species from south Brazil and taxonomic rearrangements for Taydebis Freitas, 2013 and Hermeuptychia Forster, 1964 (Lepidoptera: Nymphalidae: Satyrinae)

scholarly article published in 2021

A phylogenetic reassessment of Hyalenna Forbes and Dircenna Doubleday, with a revision of Hyalenna (Lepidoptera: Nymphalidae: Ithomiinae)

scholarly article by Keith Richard Willmott & Gerardo Lamas published 5 June 2006 in Systematic Entomology

A revision of the new genus Amiga Nakahara, Willmott & Espeland, gen. n., described for Papilio arnaca Fabricius, 1776 (Lepidoptera, Nymphalidae, Satyrinae)

scientific article published on 31 January 2019

Before it is too late: description of a new genus and species of butterfly from a highly threatened Brazilian biome

scientific article published in April 2018

Contrasting patterns of Andean diversification among three diverse clades of Neotropical clearwing butterflies.

scientific article

Contribution to the knowledge of Ecuadorian Pronophilini, Part III. Three new species and five new subspecies of Lymanopoda (Lepidoptera: Nymphalidae: Satyrinae)

scientific article published in 1999

Correlations between adult mimicry and larval host plants in ithomiine butterflies

scientific article published in August 2004

Cryptic speciation associated with geographic and ecological divergence in two Amazonian Heliconius butterflies

article by Neil Rosser et al published 4 August 2018 in Zoological Journal of the Linnean Society

DNA barcodes uncover hidden taxonomic diversity behind the variable wing patterns in the Neotropical butterfly genusZaretis(Lepidoptera: Nymphalidae: Charaxinae)

scholarly article by Fernando Maia Silva Dias et al published 1 August 2018 in Zoological Journal of the Linnean Society

Description d’une nouvelle espèce d’Euptychiina du genre Chloreuptychia (Lepidoptera, Nymphalidae, Satyrinae)

scientific article published on 19 June 2019

Description of Emeryus Zacca, Mielke & Casagrande gen. nov. (Lepidoptera: Nymphalidae) to accommodate three species formerly placed in Paryphthimoides Forster, 1964

scientific article published in August 2020

Description of a new genus and species for a common and widespread Amazonian satyrine butterfly (Lepidoptera: Nymphalidae: Satyrinae: Satyrini)

scientific article published on 04 December 2020

Description of a new species of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the western Andes

scientific article (publication date: 3 November 2016)

Diversification of clearwing butterflies with the rise of the Andes

scientific article

Fifty years without a name: a new species of Splendeuptychia Forster (Lepidoptera: Nymphalidae: Satyrinae)

scholarly article published in 2021

Five new Penaincisalia species (Lepidoptera: Lycaenidae: Eumaeini) from the Andes of southern Ecuador and northern Peru

scientific article (publication date: 2005)

Four hundred shades of brown: Higher level phylogeny of the problematic Euptychiina (Lepidoptera, Nymphalidae, Satyrinae) based on hybrid enrichment data

scientific article published on 10 November 2018

Frequency dependence shapes the adaptive landscape of imperfect Batesian mimicry.

scientific article published in April 2018

Graphita gen. nov., a New Genus for Neonympha griphe C. Felder & R. Felder, 1867 (Lepidoptera, Nymphalidae, Satyrinae).

scientific article

Hard to catch: experimental evidence supports evasive mimicry

article by Erika Páez et al published 10 March 2021 in Proceedings of the Royal Society B

Heterogeneity in predator micro-habitat use and the maintenance of Müllerian mimetic diversity

scientific article published on April 30, 2011

Incompatible Ages for Clearwing Butterflies Based on Alternative Secondary Calibrations

scientific article published on 25 May 2015

Into the Andes: multiple independent colonizations drive montane diversity in the Neotropical clearwing butterflies Godyridina

scientific article

Ithomiini butterflies (Lepidoptera: Hymphalidae) of Antioquia, Colombia

scientific article

Maintaining mimicry diversity: optimal warning colour patterns differ among microhabitats in Amazonian clearwing butterflies.

scientific article published in May 2017

Molecular phylogenetics of the neotropical butterfly subtribe Oleriina (Nymphalidae: Danainae: Ithomiini).

scientific article published on 15 January 2010

Molecular systematics of the butterfly genus Ithomia (Lepidoptera: Ithomiinae): a composite phylogenetic hypothesis based on seven genes

scientific article published on 16 December 2004

Mutualistic mimicry and filtering by altitude shape the structure of Andean butterfly communities

scientific article

North Andean origin and diversification of the largest ithomiine butterfly genus

scientific article

Phylogenetic community ecology needs to take positive interactions into account: Insights from colourful butterflies

scientific article published on January 2009

Phylogenetic relationships of ithomiine butterflies (Lepidoptera: Nymphalidae: Danainae) as implied by combined morphological and molecular data

article by Andrew V. Z. Brower et al published 31 March 2014 in Systematics and Biodiversity

Rapid diversification associated with ecological specialization in Neotropical Adelpha butterflies.

scientific article published on 20 April 2015

Redescription of Yphthimoides patricia (Hayward, 1957), with taxonomic notes on the names Euptychia saltuensis Hayward, 1962 and Yphthimoides manasses (C. Felder & R. Felder, 1867) (Nymphalidae: Satyrinae)

scientific article

Remarkable sexual dimorphism, rarity and cryptic species: a revision of the ‘aegrota species group’ of the Neotropical butterfly genus Caeruleuptychia with the description of three new species (Lepidoptera, Nymphalidae, Satyrinae)

scholarly article by Shinichi Nakahara et al published 13 April 2018 in Insect Systematics and Evolution

Revalidation of Vareuptychia Forster, 1964, description of Vanima gen. nov., and notes on Euptychia cleophes Godman amp; Salvin, 1889 (Lepidoptera: Nymphalidae: Satyrinae)

scientific article published on 29 September 2020

Revision of the poorly known Neotropical butterfly genus Zischkaia Forster, 1964 (Lepidoptera, Nymphalidae, Satyrinae), with descriptions of nine new species

article by Shinichi Nakahara et al published 13 September 2019 in European Journal of Taxonomy

Solving the cryptic diversity of the genus Manerebia Staudinger in northern Peru: description of new species and considerations on the biogeographical role of the Huancabamba Deflection (Nymphalidae: Satyrinae: Pronophilina)

scholarly article published in 2021

Strikingly variable divergence times inferred across an Amazonian butterfly 'suture zone'

scientific article (publication date: 7 December 2005)

Systematics of Hypanartia (Lepidoptera: Nymphalidae: Nymphalinae), with a test for geographical speciation mechanisms in the Andes

scholarly article by Keith Richard Willmott et al published October 2001 in Systematic Entomology

Taxonomic notes on Napeogenes from Ecuador and Colombia, with the description of ten new subspecies (Lepidoptera: Ithomiinae)

scientific article published in 2008

Taxonomic notes on Papilio ocypete Fabricius, 1776 and Papilio helle Cramer, 1779 with description of two new similar species from South America (Lepidoptera: Nymphalidae: Satyrinae)

scientific article

The phylogenetic pattern of speciation and wing pattern change in neotropical Ithomia butterflies (Lepidoptera: nymphalidae).

scientific article published on July 2006

The roles of wing color pattern and geography in the evolution of Neotropical Preponini butterflies

scientific article published on 03 October 2020

Two New Species of Taygetina With a Possible Case of ‘Juxta Loss’ in Butterflies (Lepidoptera: Nymphalidae: Satyrinae)

scholarly article by Shinichi Nakahara et al published 1 November 2019 in Insect Systematics and Diversity

Urban areas as refuges for endemic fauna: description of the immature stages of<i>Catasticta flisa duna</i>(Eitschberger &amp; T. Racheli, 1998) (Lepidoptera: Pieridae) and its ecological interactions

scientific article published in 2020

‘Species’ from two different butterfly genera combined into one: description of a new genus of Euptychiina (Nymphalidae: Satyrinae) with unusually variable wing pattern

article by André Victor Lucci Freitas et al published April 2016 in Revista Brasileira de Entomologia